Oh, this is a situation I actually had.
When you check retreive control record data from IDoc in the Communication channel, it will use that data from the EDI_DC40 field in your message mapping instead of the Sytem landscape values you have on your system. I have to do it for my tests because the servers I use are not in the SDL.
When you don't have the apply records from payload box checked, XI will automatically pull the values from your SDL, and ignore the values in EDI_DC40 from your mapping, that's why you can disable the EDI_DC40 field.
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